Vivian's family was killed in America when she was a young girl, after which she moved to Romania to live with her aunt Astrid. Astrid is a former mate of the pack's leader, Gabriel, and together they have a son named Rafe. According to pack law, the leader chooses a new mate every seven years. The choosing is a few months away and Gabriel wants Vivian as his mate, despite her reluctance. This is because of a prophecy which states that a female loup-garou will lead all loup-garou into the age of hope. Gabriel thinks this wolf will be Vivian, and for this reason he wants to marry her.
While sitting in a church one night (having broken into it) she meets a human, Aiden, who is an artist (who broke into the church for inspiration for his graphic novel series) researching the legend of the loup-garou. Aiden is instantly smitten and pursues her for several days before she finally agrees to start seeing him. They meet in secret and fall in love. They are soon discovered by Vivian's cousin Rafe and his four friends, who grew up with Vivian; throughout the book Rafe and his friends are referred to as the Five. After Rafe tell Gabriel about the relationship, he orders Rafe to do whatever is necessary to get Aiden out of the city- even if it means killing Aiden.
Rafe sends Aiden a note, pretending it is from Vivian and asking him to meet her in a chapel outside the city. After Rafe threatens him, Aiden throws him into the table where he cuts himself. Rafe's spilling of blood forces his eyes to turn gold and reveal his loup-garou side. He attempts to kill Aiden, however silver is fatal to the loup-garou and can poison them when mixes with their blood. Aiden uses his silver pentagram necklace to kill Rafe, in self-defense.
Gabriel and Astrid are devastated at the death of their son, and Aiden is caught by the other loup-garoux. Every month at full moon, the pack leads a hunt in the forest. The human prey is chosen because they are a danger to the pack or because they've offended one of its members. If the human manages to cross the river in the forest, they're allowed to live, but no one has ever managed to reach the river. Aiden is chosen to be that moon cycle's prey and he runs for his life. Fortunately, he'd stolen a silver knife from a restaurant earlier, and when two wolves corner him he is able to defend himself. He reaches the river and with great difficulty crosses it, however Gabriel is furious and breaks his own law, he follows Aiden and attacks. Vivian jumps in and she and Gabriel fight in their wolf forms; she wins and throws him into the river. Aiden attacks her, not recognizing her in her wolf skin, and stabs her with the knife. When she changes back into her human skin she reveals who she is, he is stricken with guilt and is eager and recluctant to help her.
They flee to a building where films were once processed, because it is riddled with silver dust the loup-garou would not dare enter a place so littered with silver to find them. Astrid, however, realizes where Vivian had gone and confronts them with a gun. Vivian pleads for Aiden's life, reminding Astrid of how important it is to be with someone you love, and Astrid (who is in love with Gabriel even though he is no longer her mate) lets them free. They retrieve an antidote for Vivian by threatening a loup-garou pharmacist, but the man manages to alert the pack to their presence and Vivian is captured.
In the wolves' territory, Gabriel orders one of the loup-garou to kill Vivian, but Aiden jumps in with a gun full of silver bullets and a massive fight begins. During the shooting, several kegs of alcohol shatter and their contents spill onto the floor and Aiden sets the building on fire. Vivian aims a gun at Gabriel, but he blackmails her by telling her she how she is low enough to try to kill one of her own kind in order to defend a human. Aiden pleads with Vivian to shoot Gabriel, to which she replies, "I can't." When Gabriel changes into his wolf form and goes to attack Aiden, Vivian shoots him. Vivian feels guilty for having killed Gabriel, as tears roll down her cheeks and she strokes Gabriel's fur. She helps the rest of the Loup-garou escape from the fire and she and Aiden run away together.
The last scene is of them sitting in Gabriel's car and driving out of the city, on their way to Paris. As they drive, several people in the streets bow their heads and smile at the car, as they believe it to be the pack leader Gabriel. These people are loup-garoux and they have no knowledge that Gabriel is dead. This was the only way for Vivian and Aiden to escape the city undetected.
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